Cronin DT, Sesink Clee PR, Mitchell MW, et al. Conservation strategies for understanding and combating the primate bushmeat trade on Bioko Island, Equatorial Guinea. Am J Primatol. 2017;79:e22663. (LINK).


Primate communities along a protected area border: the value of a long-term research site to primate conservation. Forrest, D.L, F. Muatiche, M.K. Gonder, D.T. Cronin, , African Primates, 2017, 12: 23-26.



S. Heritage, D. Fernández, H.M. Sallam, D.T. Cronin, J.M. Esara Echube, and E.R. Seiffert. Ancient phylogenetic divergence of the enigmatic African rodent Zenkerella and the origin of anomalurid gliding.  PeerJ, 4: e2320. (LINK).


D.T. Cronin, C. Riaco, J.M. Linder, R.A. Bergl, M.K. Gonder, M.P. O'Connor, and G.W. Hearn. Impact of gun-hunting on monkey species and implications for primate conservation on Bioko Island, Equatorial Guinea. Biological Conservation, 197: 180-189. (LINK).



Hoffman, M., D.T. Cronin, G.W. Hearn, T.M. Butynski, and E. Do Linh San. 2015. A review of evidence for the presence of Two-spotted Palm Civet Nandinia binotata and four other small carnivores on Bioko, Equatorial Guinea. Small Carnivore Conservation 52 & 53: 13-23. (LINK)


Evans, B.J., T.F. Carter, E. Greenbaum, V. Gvoždík, D.B. Kelley, P.J. McLaughlin, et al. 2015. Genetics, Morphology, Advertisement Calls, and Historical Records Distinguish Six New Polyploid Species of African Clawed Frog (Xenopus, Pipidae) from West and Central Africa. PLoS ONE 10(12): e0142823. doi:10.1371/journal.pone.0142823 (LINK)


Owens, J.R., S. Honarvar, M. Nessel, and G.W. Hearn. From frugivore to folivore: Altitudinal variations in the diet and feeding ecology of the Bioko Island drill (Mandrillus leucophaeus poensis). American Journal of Primatology 77(12):1263-1275. (LINK)


Cronin, D.T., S. Woloszynek, W.A. Morra, S. Honarvar, J.M. Linder, M.K. Gonder, M.P. O'Connor, and G.W. Hearn. Long-term urban market dynamics reveal increased bushmeat carcass volume despite economic growth and proactive environmental legislation on Bioko Island, Equatorial Guinea. PLoS ONE 10(7): e0134464. doi:10.1371/journal.pone.0134464 (LINK)


Cronin, D.T., D. Bocuma Meñe, C. Perella, D. Fernández, G.W. Hearn, and M.K. Gonder. 2015. The Future of the Biodiversity of the Gran Caldera Scientific Reserve: Translating Science into Policy to Develop an Effective Management Plan for the Gran Caldera Scientific Reserve. Report prepared for the Symposium on the Future of the Biodiversity of the Gran Caldera Scientific Reserve by the Bioko Biodiversity Protection Program. Version 1.0, Philadelphia, 42 pages. (ENGLISH) (SPANISH)



Cronin, D.T., M.B. Libalah, R.A. Bergl, and G.W. Hearn. 2014. Biodiversity and conservation of tropical montane ecosystems in the Gulf of Guinea, West Africa. Arctic, Antarctic, and Alpine Research 46(4):891-904. (PDF)


Cronin, D. T., G.W. Hearn, and J.F. Oates. 2014. Bioko red colobus Piliocolobus pennantii pennantii (Waterhouse, 1838). In: C. Schwitzer, R. A. Mittermeier, A. B. Rylands, L. A. Taylor, F. Chiozza, E. A. Williamson, J. Wallis and F. E. Clark (eds.), Primates in Peril: The World’s 25 Most Endangered Primates 2012-2014, pp. 17-19. IUCN SSC Primate Specialist Group (PSG), International Primatological Society (IPS), Conservation International (CI), and Bristol Zoological Society, Arlington, VA.(PDF)


Umile, T.P., P.J. McLaughlin, K.R. Johnson, S. Honarvar, A.L. Blackman, E.A. Burzynski, R.W. Davis, T.L. Teotonio, G.W. Hearn, C.A. Hughey, A.F. Lagalante, K.P.C. Minbiole. 2014. Nonlethal Amphibian Skin Swabbing of Cutaneous Natural Products for HPLC Fingerprinting. Analytical Methods. 6: 3277-3284. (PDF)


Eberhard, M.L., J.R. Owens, H.S. Bishop, M.E. de Almeida, A.J. de Silva, G.W. Hearn, S. Honarvar. 12014. Cyclospora from Drills (Mandrillus leucophaeus poensis) on Bioko Island, Equatorial Guinea. Emerging Infectious Diseases 20 (3): 510-511. (PDF).


Cronin, D.T., J.R. Owens, H. Choi, S. Hromada, R. Malhotra, F. Roser, R.A. Bergl. 2014. Where has all our research gone? A 20-year assessment of the peer-reviewed wildlife conservation literature. International Journal of Comparative Psychology 27(1): 101-116. (PDF)



Cronin, D.T., C. Riaco, G.W. Hearn. 2013. Survey of Threatened Monkeys in the Iladyi River Valley Region, Southeastern Bioko Island, Equatorial Guinea. African Primates 8: 1-8. (PDF)



Ting, N., C. Astaras, G. Hearn, S. Honarvar, J. Corush, A. Burrell, N. Phillips, B. Morgan, E. Gadsby, R. Raaum, C. Roos. 2012. Genetic signatures of a demographic collapse in a large-bodied forest dwelling primate (Mandrillus leucophaeus). Ecology and Evolution 2: 550-561.(PDF)



Fitzgerald, D. B., E. Ordway, S. Honarvar, G. W. Hearn. 2011. Challenges confronting Sea Turtle Conservation on Bioko Island, Equatorial Guinea. Chelonian Conservation and Biology. Chelonian Conservation and Biology 10:177-180.(PDF)


Honarvar, S., M. C. Brodsky, D. B. Fitzgerald, K. L. Rosenthal and G. W. Hearn . 2011. Changes in plasma chemistry and reproductive output of nesting leatherbacks. Herpetologica 67:222-235.(PDF)



Cronin, D. T., D. Bocuma Meñe, T. B. Butynski, J. M. E. Echube, G. W. Hearn, S. Honarvar, J. R. Owens, and C. P. Bohome. 2010. Opportunities Lost: The Rapidly Deteriorating Conservation Status of the Monkeys on Bioko Island, Equatorial Guinea (2010). A Report to the Government of Equatorial Guinea by the Bioko Biodiversity Protection Program, Drexel University, Philadelphia, Pennsylvania.(PDF)


Worobey, M., P. Telfer, S. Souquiere, M. Hunter, C. A. Coleman, M. J. Metzger, P. Reed, M. Makuwa, G. Hearn, S. Honarvar, P. Roques, C. Apetrei, M. Kazanji, and P. A. Marx. 2010. Island Biogeography Reveals the Deep History of SIV. Science 329:1487-1487.(PDF)



Butynski, T. B., Y. A. de Jong, and G. W. Hearn. 2009. Body Measurements for the Monkeys of Bioko Island, Equatorial Guinea. Primate Conservation 24:99-105.(PDF)


Morra, W., G. Hearn, and A. J. Buck. 2009. The market for bushmeat: Colobus Satanas on Bioko Island. Ecological Economics 68:2619-2626.(PDF)


Witherington,B., K. Pendoley, G.W. Hearn, and S. Honarvar. 2009. Ancient mariners, ancient fuels: How sea turtles cope with our modern fossil fuel dependency. The State of the World Sea Turtles Report 4:38-41.



Hearn, G. W., W. Morra, and T. B. Butynski. 2006. Monkeys in Trouble: The Rapidly Deteriorating Conservation Status of the Monkeys on Bioko Island, Equatorial Guinea. Bioko Biodiversity Protection Program, Arcadia University, Glenside, Pennsylvania.(PDF)


Rader, H., M.A. Ela Mba, W.A. Morra, and G.W. Hearn. 2006. Marine Turtles on the Southern Coast of Bioko Island (Gulf of Guinea, Africa) 2001-2005. Marine Turtle Newsletter 111:8-10.(PDF)


All photos are credited to National Geographic Photographers Tim Laman, Ian Nichols, Joel Sartore, and Christian Ziegler, as well as numerous members of BBPP (staff, students, and volunteers).