meet our primates









Red Colobus


Crowned Guenon



Binomial: Mandrillus leucophaeus poensis

Common: Drill



Baboon-like with a prominent muzzle, a black face, and a very short tail. Mostly gray-brown coat, with lighter ventral surfaces and a yellow-brown crown. Hair on the belly midline is long and white, seen as a prominent hanging fringe in males. Males are distinguished by their rump and genitalia, which are bright blue, mauve, pink, and red, and scarlet inner thighs. Female coloration is drab. Males also possess large canine teeth and long hair on the neck and shoulders, which forms a “mane."

Binomial: Colobus satanas satanas

Common: Black Colobus



Entirely black. Coat is long, especially on the shoulders and forehead. Has a small head, relatively long legs, and a large belly.


Binomial: Cercopithecus erythrotis erythrotis

Common: Red-eared Guenon



Rusty red colored fringe of hair on the outer margin of the ears. Tail is similar red color on ventral side, but more brown underneath with a black tip. The crown of the head, back, and outer legs are gray, with orange-brown speckles. Lower arms are dark gray, and belly is a lighter gray. The bridge of the nose has a diamond-shaped patch of red hair, the upper cheeks have tufts of white and yellow hair, and the lower cheeks have tufts of black hair.

Binomial: Cercopithecus nictitans martini

Common: Putty-nosed Guenon



The most characteristic feature is a bulbous, white nose. Crown of head, shoulders, back, and outer legs are dark gray, speckled with olive-yellow. The fore-chest is light gray. The outer arms, distal part of tail, and hands and feet are black.


Binomial: Procolobus pennantii pennantii

Common: Pennant's Red Colobus



Primarily dark charcoal gray-brown on crown of head extending over back to upper thighs. Rich orange-rust red on flanks and outer surface of limbs. Creamy white throat, underside, and inner limbs. Tail is a dark reddish brown. Face, hands, and feet extending to above the ankles are black.


Binomial: Cercopithecus pogonias pogonias

Common: Golden-bellied Crowned Guenon



Distinguished by a crest of black hair on the crown of its head and bright gold

hair on the chest, belly, and inner limbs. Outer surfaces of limbs are fawn or gray. A dorsal band of dark hair extends from the shoulders to the root of the tail, widening at the posterior. Tail is dark on the dorsal side and golden-brown on the ventral side.

Binomial: Allochrocebus preussi insularis

Common: Preuss's Monkey



The crown, shoulders, flanks, thighs, and mid-tail are dark gray with silvery speckles. The throat is silvery white and the arms, hands, feet, and tip of the tail are black. The middle back is red-brown, and the face and underside are gray.

Our work

bushmeat study

The hunting of wildlife for human consumption, known as bushmeat in Africa, is a luxury market on Bioko Island. Bushmeat consumed as a delicacy can range from tiny blue duikers to endangered monkeys and massive marine turtles. Because bushmeat is relatively scarce, and growing scarcer, it is priced well above other protein sources such as fish, beef, or chicken. With increasing globalization and accessibility, the market for bushmeat grows, enticing more local people to hunt wildlife for profit.


Since 1997, BBPP has conducted daily surveys of animals for sale at the Malabo bushmeat market. Historically rising numbers of carcasses in the market have begun to drop since late 2010, indicating instability in the Bioko Island economy and the possible success of recent conservation efforts. BBPP research focuses on understanding the drivers and significant periods of intervention in the Malabo market series.


In 2007, the government of Equatorial Guinea instituted a ban on primate hunting and trade. Unfortunately, the ban was not enforced, and was followed by an increase in bushmeat hunting. The results from our ongoing bushmeat study emphasize the negative impact that rapid development and unenforced legislation have had on Bioko’s wildlife, and demonstrate the need for governmental support to implement conservation strategies meant to prevent extinctions of tropical wildlife.


Carcasses Counted to Date:


Days Counted to Date:

Bioko microbiome project

Primates and their microbiomes share a long evolutionary history. Throughout primate evolution, gut microbiota have been important in defining critical metabolic pathways that have given rise to unique dietary adaptations because they facilitate host digestion, metabolism, production of vitamins, and immune response. Increasingly, studies have revealed that the bacterial communities in primate gastrointestinal tracts are affected by a variety of factors (e.g. diet and habitat disturbance), but the external factors that influence composition of wild primate microbiomes and the abundance of specific microbial species remain controversial (Amato et al. 2013; Gomez et al. 2015). In addition, Bioko Island has diverse flora, fauna, and climatic conditions across the elevational gradient that could affect the gut flora of these species, thereby potentially offering a rich system to examine the factors that influence the diversity of the gut microbiome of non-human primate communities (McCord et al. 2014). The goal of this study is to describe the gut microbial diversity of all non-human primates on Bioko Island, determine what factors are influencing the gut composition, and what roles microbiota play in niche competition between these closely related primates.


For more information please contact Bryan Featherstone, Ph.D. Student (

All photos are credited to National Geographic Photographers Tim Laman, Ian Nichols, Joel Sartore, and Christian Ziegler, as well as numerous members of BBPP (staff, students, and volunteers).